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The snake uses there tongue totally differently to the way mammals do. Mammals use their tongues to eat with, and the tongue is an extremely powerful muscle, so having a single solid bulk gives a stronger muscle.The snake uses its tongue as a chemical detector, much as insects use their antenna, and like an insects antenna, having it face two directions gives it the capability to have some slight directionality in detecting smell (i.e. if the left part of the tongue tastes a smell stronger then the right part, then it can deduce that the thing it is smelling is more to the left than to the right).
I think the ears would have been better to detect prey. With tongue the directional presence is highly dependent on air flow. First the prey is difficult to catch and then you have this dependency on wind. And this extra assumption of tongue replacing nose and ears.Insignificant observation I would say.Moisture content of the atmospehere and direction of wind can be determined using tongue. Snakes like to move towards colder regions when hot and towards hotter regions when cold..And importantly it appears to me that forked tongue offers more refined sense of taste. They taste poisons as well. Humans or other animals do not try to eat animals which are poisnous.. but these reptiles do eat poisnous insects animals. And it is important for them to distinguish the extended list of "tastes".I think poisnous spider must also be having some mechanism to distinguish different dishes.