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If both branes exist within the bulk it seems to me that the branes must share some dimensions of the bulk, like a plane intersecting a solid. If this is so, and even if we assume that the bulk is non-homogeneous i.e. not smooth, then as the two branes approach each other they should become more and more alike.
Also, at some point the approaching brane must pass through the gravity brane (unless it's always been between us), at which point it should have been identical to the gravity brane.
and then begs the question: how and why is the other brane different to ours, if it's only just further back down the bulk in a region that we have already passed through?
Particles & forces could be localised or sequestered. They could either be confined to the brane on which they originate, or could not travel far from it, and, in the case of localisation, the strength of the forces would decrease exponentially as you moved away.