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BTW, mitochondria have molecules all over their surface that spin constantly at about 100 to 200 revolutions per second. (Now I know why my ears ring constantly.)
How far back did this symbiotic relationship go ...
The oldest known fossilized prokaryotes were laid down approximately 3.5 billion years ago, only about 1 billion years after the formation of the Earth's crust... Eukaryotes only appear in the fossil record later, and may have formed from endosymbiosis of multiple prokaryote ancestors. The oldest known fossil eukaryotes are about 1.7 billion years old. However, some genetic evidence suggests eukaryotes appeared as early as 3 billion years ago.
That fact that we functioned without them is boggling.